^ Alpha Centauri is the fifth founding member of the Federation, but I don't know what area of expertise its sub-fleet (if it had one) could possibly be responsible for. You'll have to ask @Christopher about that one.
Although whatever sub-fleet that might be, the Defiant (NCC-1764) was probably part of it, as its crew (as shown in "In A Mirror, Darkly") wore an insignia that was different from all four of the other sub-fleets I mentioned. Again, though, I don't know what area of Starfleet this could possibly refer to. Maybe @Christopher's saving that for a future novel.
Although whatever sub-fleet that might be, the Defiant (NCC-1764) was probably part of it, as its crew (as shown in "In A Mirror, Darkly") wore an insignia that was different from all four of the other sub-fleets I mentioned. Again, though, I don't know what area of Starfleet this could possibly refer to. Maybe @Christopher's saving that for a future novel.

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