Therin of Andor wrote:
It's possible the cel painters had not even seen TOS, let alone have access to stills of the show. Remember the Gold Key comic artists had also never seen the show.
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Alberto Giolitti never screened the series, but he worked from a stack of publicity photos from seasons one and two, which is why some famous poses of the series leads and
Enterprise can be found among the line art.
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Were Filmation's colourists even based in USA?
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At the time of TAS' production, yes.